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a γH2AX foci (DSBs) original positions of foci repair complexes foci movement danger of translocation or unreparable DSB bc IR Position First Hypothesis Breakage First Hypothesis in combination with repair factories de Eu C Het B B A D ACD Decondensation model of formation of chromosomal translocations In the case of the Position First Hypothesis, DSBs (circles) induced by ionizing radiation (a) start to colocalize with repair proteins (white-black diamonds) and are repaired at the sites of their origin (b); translocations, therefore, form only between loci that have been close to each other already before their damage (b).
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